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We intuitively say that a plant is stressed when there is a reduction in some physiological rate (water or nutrient absorption, photosynthesis, respiration, growth, development, reproduction or others) below the maximum possible rate expressed under optimal conditions (Salisbury and Ross, 1992; Lambers et al. 1998), thus not attaining its genotypic potential. There are many environmental stress factors that can limit growth and development; Levitt (1980) proposed to classify them in biotic and abiotic factors (Table 1), causing biotic or abiotic stress in plants, respectively.