Department of Zoology, University of Cambridge

University of Liverpool

Professor of Biology, University of St Andrews

Professor of Comparative Psychology, Cambridge University

Edward Grey Institute, Department of Zoology, University of Oxford

University of Bristol

Professor emeritus, Animal Ecology, University of Gothenburg

Humboldt University and Leibniz-Institute of Freshwater Ecology and Inland Fisheries

University of British Columbia

Department of Veterinary Medicine, University of Cambridge